In 1876–77, an England team took part in what was retrospectively recognised as the first-ever Test match at the Melbourne Cricket Ground against Australia. The rivalry between England and Australia gave birth to The Ashes in 1882, and this has remained Test cricket's most famous contest. Test cricket began to expand in 1888–89 when South Africa played England.
The phylogenetic relationships of the Gryllidae, summarized by Darryl Gwynne in 1995 from his own work (using mainly anatomical characteristics) and that of earlier authors,[a] are shown in the following cladogram, with the Orthoptera divided into two main groups, Ensifera (crickets sensu lato) and Caelifera (grasshoppers). Fossil Ensifera are found from the late Carboniferous period (300 Mya) onwards, and the true crickets, Gryllidae, from the Triassic period (250 to 200 Mya).
More than 900 species of crickets are described; the Gryllidae are distributed all around the world except at latitudes 55° or higher, with the greatest diversity being in the tropics. They occur in varied habitats from grassland, bushes, and forests to marshes, beaches, and caves. Crickets are mainly nocturnal, and are best known for the loud, persistent, chirping song of males trying to attract females, although some species are mute. The singing species have good hearing, via the tympana on the tibiae of the front legs.
The innings (ending with 's' in both singular and plural form) is the term used for each phase of play during a match. Depending on the type of match being played, each team has either one or two innings. Sometimes all eleven members of the batting side take a turn to bat but, for various reasons, an innings can end before they have all done so. The innings terminates if the batting team is "all out", a term defined by the Laws: "at the fall of a wicket or the retirement of a batsman, further balls remain to be bowled but no further batsman is available to come in". In this situation, one of the batsmen has not been dismissed and is termed not out; this is because he has no partners left and there must always be two active batsmen while the innings is in progress.
Other defensive strategies are the use of camouflage, fleeing, and aggression. Some species have adopted colourings, shapes, and patterns that make it difficult for predators that hunt by sight to detect them. They tend to be dull shades of brown, grey, and green that blend into their background, and desert species tend to be pale. Some species can fly, but the mode of flight tends to be clumsy, so the most usual response to danger is to scuttle away to find a hiding place.
If a fielder is injured or becomes ill during a match, a substitute is allowed to field instead of him, but the substitute cannot bowl or act as a captain. The substitute leaves the field when the injured player is fit to return. The Laws of Cricket were updated in 2017 to allow substitutes to act as wicket-keepers, a situation that first occurred when Mumbai Indians' wicket-keeper Ishan Kishan was injured in a match on 18 April 2018.
The bat is made of wood, usually salix alba (white willow), and has the shape of a blade topped by a cylindrical handle. The blade must not be more than four and one quarter inches (108 mm) wide and the total length of the bat not more than 38 inches (965 mm). There is no standard for the weight, which is usually between 2 lb 7 oz and 3 lb (1.1 and 1.4 kg).
Some species of cricket are polyandrous. In Gryllus bimaculatus, the females select and mate with multiple viable sperm donors, preferring novel mates. Female Teleogryllus oceanicus crickets from natural populations similarly mate and store sperm from multiple males. Female crickets exert a postcopulatory fertilization bias in favour of unrelated males to avoid the genetic consequences of inbreeding. Fertilization bias depends on the control of sperm transport to the sperm storage organs. The inhibition of sperm storage by female crickets can act as a form of cryptic female choice to avoid the severe negative effects of inbreeding. Controlled-breeding experiments with the cricket Gryllus firmus demonstrated inbreeding depression, as nymphal weight and early fecundity declined substantially over the generations' this was caused as expected by an increased frequency of homozygous combinations of deleterious recessive alleles.
Cricket is one of many games in the "club ball" sphere that basically involve hitting a ball with a hand-held implement; others include baseball, golf, hockey, tennis, squash, badminton and table tennis. In cricket's case, a key difference is the existence of a solid target structure, the wicket (originally, it is thought, a "wicket gate" through which sheep were herded), that the batsman must defend. The cricket historian Harry Altham identified three "groups" of "club ball" games: the "hockey group", in which the ball is driven to and fro between two targets (the goals); the "golf group", in which the ball is driven towards an undefended target (the hole); and the "cricket group", in which "the ball is aimed at a mark (the wicket) and driven away from it".
Although the main object of the game has always been to score the most runs, the early form of cricket differed from the modern game in certain key technical aspects. The ball was bowled underarm by the bowler and all along the ground towards a batsman armed with a bat that, in shape, resembled a hockey stick; the batsman defended a low, two-stump wicket; and runs were called "notches" because the scorers recorded them by notching tally sticks.
At the grassroots level, local club cricket is essentially an amateur pastime for those involved but still usually involves teams playing in competitions at weekends or in the evening. Schools cricket, first known in southern England in the 17th century, has a similar scenario and both are widely played in the countries where cricket is popular. Although there can be variations in game format, compared with professional cricket, the Laws are always observed and club/school matches are therefore formal and competitive events. The sport has numerous informal variants such as French cricket.