A phylogenetic study by Jost & Shaw in 2006 using sequences from 18S, 28S, and 16S rRNA supported the monophyly of Ensifera. Most ensiferan families were also found to be monophyletic, and the superfamily Gryllacridoidea was found to include Stenopelmatidae, Anostostomatidae, Gryllacrididae and Lezina. Schizodactylidae and Grylloidea were shown to be sister taxa, and Rhaphidophoridae and Tettigoniidae were found to be more closely related to Grylloidea than had previously been thought. The authors stated that "a high degree of conflict exists between the molecular and morphological data, possibly indicating that much homoplasy is present in Ensifera, particularly in acoustic structures." They considered that tegmen stridulation and tibial tympanae are ancestral to Ensifera and have been lost on multiple occasions, especially within the Gryllidae.
Crickets have a cosmopolitan distribution, being found in all parts of the world with the exception of cold regions at latitudes higher than about 55° North and South. They have colonised many large and small islands, sometimes flying over the sea to reach these locations, or perhaps conveyed on floating timber or by human activity. The greatest diversity occurs in tropical locations, such as in Malaysia, where 88 species were heard chirping from a single location near Kuala Lumpur. A greater number than this could have been present because some species are mute.
The decision to attempt a run is ideally made by the batsman who has the better view of the ball's progress, and this is communicated by calling: usually "yes", "no" or "wait". More than one run can be scored from a single hit: hits worth one to three runs are common, but the size of the field is such that it is usually difficult to run four or more. To compensate for this, hits that reach the boundary of the field are automatically awarded four runs if the ball touches the ground en route to the boundary or six runs if the ball clears the boundary without touching the ground within the boundary. In these cases the batsmen do not need to run. Hits for five are unusual and generally rely on the help of "overthrows" by a fielder returning the ball. If an odd number of runs is scored by the striker, the two batsmen have changed ends, and the one who was non-striker is now the striker. Only the striker can score individual runs, but all runs are added to the team's total.
Captive crickets are omnivorous; when deprived of their natural diet, they accept a wide range of organic foodstuffs. Some species are completely herbivorous, feeding on flowers, fruit, and leaves, with ground-based species consuming seedlings, grasses, pieces of leaf, and the shoots of young plants. Others are more predatory and include in their diet invertebrate eggs, larvae, pupae, moulting insects, scale insects, and aphids. Many are scavengers and consume various organic remains, decaying plants, seedlings, and fungi. In captivity, many species have been successfully reared on a diet of ground, commercial dry dog food, supplemented with lettuce and aphids.