The phylogenetic relationships of the Gryllidae, summarized by Darryl Gwynne in 1995 from his own work (using mainly anatomical characteristics) and that of earlier authors,[a] are shown in the following cladogram, with the Orthoptera divided into two main groups, Ensifera (crickets sensu lato) and Caelifera (grasshoppers). Fossil Ensifera are found from the late Carboniferous period (300 Mya) onwards, and the true crickets, Gryllidae, from the Triassic period (250 to 200 Mya).
According to the social historian Derek Birley, there was a "great upsurge of sport after the Restoration" in 1660. Gambling on sport became a problem significant enough for Parliament to pass the 1664 Gambling Act, limiting stakes to £100 which was, in any case, a colossal sum exceeding the annual income of 99% of the population. Along with prizefighting, horse racing and blood sports, cricket was perceived to be a gambling sport. Rich patrons made matches for high stakes, forming teams in which they engaged the first professional players. By the end of the century, cricket had developed into a major sport which was spreading throughout England and was already being taken abroad by English mariners and colonisers – the earliest reference to cricket overseas is dated 1676. A 1697 newspaper report survives of "a great cricket match" played in Sussex "for fifty guineas apiece" – this is the earliest known match that is generally considered top-class.
Females are generally attracted to males by their calls, though in nonstridulatory species, some other mechanism must be involved. After the pair has made antennal contact, a courtship period may occur during which the character of the call changes. The female mounts the male and a single spermatophore is transferred to the external genitalia of the female. Sperm flows from this into the female's oviduct over a period of a few minutes or up to an hour, depending on species. After copulation, the female may remove or eat the spermatophore; males may attempt to prevent this with various ritualised behaviours. The female may mate on several occasions with different males.
A match's statistics are summarised on a scorecard. Prior to the popularisation of scorecards, most scoring was done by men sitting on vantage points cuttings notches on tally sticks and runs were originally called notches. According to Rowland Bowen, the earliest known scorecard templates were introduced in 1776 by T. Pratt of Sevenoaks and soon came into general use. It is believed that scorecards were printed and sold at Lord's for the first time in 1846.
Other defensive strategies are the use of camouflage, fleeing, and aggression. Some species have adopted colourings, shapes, and patterns that make it difficult for predators that hunt by sight to detect them. They tend to be dull shades of brown, grey, and green that blend into their background, and desert species tend to be pale. Some species can fly, but the mode of flight tends to be clumsy, so the most usual response to danger is to scuttle away to find a hiding place.
The innings (ending with 's' in both singular and plural form) is the term used for each phase of play during a match. Depending on the type of match being played, each team has either one or two innings. Sometimes all eleven members of the batting side take a turn to bat but, for various reasons, an innings can end before they have all done so. The innings terminates if the batting team is "all out", a term defined by the Laws: "at the fall of a wicket or the retirement of a batsman, further balls remain to be bowled but no further batsman is available to come in". In this situation, one of the batsmen has not been dismissed and is termed not out; this is because he has no partners left and there must always be two active batsmen while the innings is in progress.
In 1876–77, an England team took part in what was retrospectively recognised as the first-ever Test match at the Melbourne Cricket Ground against Australia. The rivalry between England and Australia gave birth to The Ashes in 1882, and this has remained Test cricket's most famous contest. Test cricket began to expand in 1888–89 when South Africa played England.
Cricket has close historical ties with Australian rules football and many players have competed at top levels in both sports. In 1858, prominent Australian cricketer Tom Wills called for the formation of a "foot-ball club" with "a code of laws" to keep cricketers fit during the off-season. The Melbourne Football Club was founded the following year, and Wills and three other members codified the first laws of the game. It is typically played on modified cricket fields.
Male crickets establish their dominance over each other by aggression. They start by lashing each other with their antennae and flaring their mandibles. Unless one retreats at this stage, they resort to grappling, at the same time each emitting calls that are quite unlike those uttered in other circumstances. When one achieves dominance, it sings loudly, while the loser remains silent.
The batsman on strike (i.e. the "striker") must prevent the ball hitting the wicket, and try to score runs by hitting the ball with his bat so that he and his partner have time to run from one end of the pitch to the other before the fielding side can return the ball. To register a run, both runners must touch the ground behind the popping crease with either their bats or their bodies (the batsmen carry their bats as they run). Each completed run increments the score of both the team and the striker.
Crickets often appear as characters in literature. The Talking Cricket features in Carlo Collodi's 1883 children's book, The Adventures of Pinocchio, and in films based on the book. The eponymous insect is central to Charles Dickens's 1845 The Cricket on the Hearth, as is the chirping insect in George Selden's 1960 The Cricket in Times Square. Crickets are celebrated in poems by William Wordsworth, John Keats, and Du Fu. They are kept as pets in countries from China to Europe, sometimes for cricket fighting. Crickets are efficient at converting their food into body mass, making them a candidate for food production. They are used as human food in Southeast Asia, where they are sold deep-fried in markets as snacks. They are also used to feed carnivorous pets and zoo animals. In Brazilian folklore, crickets feature as omens of various events.
Limited overs cricket is always scheduled for completion in a single day. There are two types: List A which normally allows fifty overs per team; and Twenty20 in which the teams have twenty overs each. Both of the limited overs forms are played internationally as Limited Overs Internationals (LOI) and Twenty20 Internationals (T20I). List A was introduced in England in the 1963 season as a knockout cup contested by the first-class county clubs. In 1969, a national league competition was established. The concept was gradually introduced to the other leading cricket countries and the first limited overs international was played in 1971. In 1975, the first Cricket World Cup took place in England. Twenty20 is a new variant of limited overs itself with the purpose being to complete the match within about three hours, usually in an evening session. The first Twenty20 World Championship was held in 2007. Limited overs matches cannot be drawn, although a tie is possible and an unfinished match is a "no result".
In 1975, Dr. William H. Cade discovered that the parasitic tachinid fly Ormia ochracea is attracted to the song of the cricket, and uses it to locate the male to deposit her larvae on him. It was the first known example of a natural enemy that locates its host or prey using the mating signal. Since then, many species of crickets have been found to be carrying the same parasitic fly, or related species. In response to this selective pressure, a mutation leaving males unable to chirp was observed amongst a population of Teleogryllus oceanicus on the Hawaiian island of Kauai, enabling these crickets to elude their parasitoid predators. A different mutation with the same effect was also discovered on the neighboring island of Oahu (ca. 100 miles (160 km) away). Recently, new "purring" males of the same species in Hawaii are able to produce a novel auditory sexual signal that can be used to attract females while greatly reducing the likelihood of parasitoid attack from the fly.
Some species, such as the ground crickets (Nemobiinae), are wingless; others have small fore wings and no hind wings (Copholandrevus), others lack hind wings and have shortened fore wings in females only, while others are macropterous, with the hind wings longer than the fore wings. In Teleogryllus, the proportion of macropterous individuals varies from very low to 100%. Probably, most species with hind wings longer than fore wings engage in flight.
Cricket flour may be used as an additive to consumer foods such as pasta, bread, crackers, and cookies. The cricket flour is being used in protein bars, pet foods, livestock feed, nutraceuticals, and other industrial uses. The United Nations says the use of insect protein, such as cricket flour, could be critical in feeding the growing population of the planet while being less damaging to the environment.
Crickets are small to medium-sized insects with mostly cylindrical, somewhat vertically flattened bodies. The head is spherical with long slender antennae arising from cone-shaped scapes (first segments) and just behind these are two large compound eyes. On the forehead are three ocelli (simple eyes). The pronotum (first thoracic segment) is trapezoidal in shape, robust, and well-sclerotinized. It is smooth and has neither dorsal or lateral keels (ridges).
Off the field in televised matches, there is usually a third umpire who can make decisions on certain incidents with the aid of video evidence. The third umpire is mandatory under the playing conditions for Test and Limited Overs International matches played between two ICC full member countries. These matches also have a match referee whose job is to ensure that play is within the Laws and the spirit of the game.
Some species, such as Gryllus assimilis, take off, fly, and land efficiently and well, while other species are clumsy fliers. In some species, the hind wings are shed, leaving wing stumps, usually after dispersal of the insect by flight. In other species, they may be pulled off and consumed by the cricket itself or by another individual, probably providing a nutritional boost.
The world's earliest known cricket match was a village cricket meeting in Kent which has been deduced from a 1640 court case recording a "cricketing" of "the Weald and the Upland" versus "the Chalk Hill" at Chevening "about thirty years since" (i.e., c. 1611). Inter-parish contests became popular in the first half of the 17th century and continued to develop through the 18th with the first local leagues being founded in the second half of the 19th.
The entomopathogenic fungus Metarhizium anisopliae attacks and kills crickets and has been used as the basis of control in pest populations. The insects are also affected by the cricket paralysis virus, which has caused high levels of fatalities in cricket-rearing facilities. Other fatal diseases that have been identified in mass-rearing establishments include Rickettsia and three further viruses. The diseases may spread more rapidly if the crickets become cannibalistic and eat the corpses.