Some species of cricket are polyandrous. In Gryllus bimaculatus, the females select and mate with multiple viable sperm donors, preferring novel mates.[21] Female Teleogryllus oceanicus crickets from natural populations similarly mate and store sperm from multiple males.[22] Female crickets exert a postcopulatory fertilization bias in favour of unrelated males to avoid the genetic consequences of inbreeding. Fertilization bias depends on the control of sperm transport to the sperm storage organs. The inhibition of sperm storage by female crickets can act as a form of cryptic female choice to avoid the severe negative effects of inbreeding.[23] Controlled-breeding experiments with the cricket Gryllus firmus demonstrated inbreeding depression, as nymphal weight and early fecundity declined substantially over the generations'[24] this was caused as expected by an increased frequency of homozygous combinations of deleterious recessive alleles.[24][25]
Crickets have a cosmopolitan distribution, being found in all parts of the world with the exception of cold regions at latitudes higher than about 55° North and South. They have colonised many large and small islands, sometimes flying over the sea to reach these locations, or perhaps conveyed on floating timber or by human activity. The greatest diversity occurs in tropical locations, such as in Malaysia, where 88 species were heard chirping from a single location near Kuala Lumpur. A greater number than this could have been present because some species are mute.[1]
By the end of the 20th century the sound of chirping crickets came to represent quietude in literature, theatre and film. From this sentiment arose expressions equating "crickets" with silence altogether, particularly when a group of assembled people makes no noise. These expressions have grown from the more descriptive, "so quiet that you can hear crickets," to simply saying , "crickets" as shorthand for "complete silence."[62]
While the umpire (1) in shot stands at the bowler's end of the pitch, his colleague stands in the outfield, usually in or near the fielding position called "square leg", so that he is in line with the popping crease (7) at the striker's end of the pitch. The bowling crease (not numbered) is the one on which the wicket is located between the return creases (12). The bowler (4) intends to hit the wicket (9) with the ball (5) or, at least, to prevent the striker (8) from scoring runs. The striker (8) intends, by using his bat, to defend his wicket and, if possible, to hit the ball away from the pitch in order to score runs.
At the tip of the abdomen is a pair of long cerci (paired appendages on rearmost segment), and in females, the ovipositor is cylindrical, long and narrow, smooth and shiny. The femora (third segments) of the back pair of legs are greatly enlarged for jumping. The tibiae (fourth segments) of the hind legs are armed with a number of moveable spurs, the arrangement of which is characteristic of each species. The tibiae of the front legs bear one or more tympani which are used for the reception of sound.[3]
If a fielder is injured or becomes ill during a match, a substitute is allowed to field instead of him, but the substitute cannot bowl or act as a captain. The substitute leaves the field when the injured player is fit to return.[74] The Laws of Cricket were updated in 2017 to allow substitutes to act as wicket-keepers,[75] a situation that first occurred when Mumbai Indians' wicket-keeper Ishan Kishan was injured in a match on 18 April 2018.[76]

One possible source for the sport's name is the Old English word "cryce" (or "cricc") meaning a crutch or staff. In Samuel Johnson's Dictionary, he derived cricket from "cryce, Saxon, a stick".[5] In Old French, the word "criquet" seems to have meant a kind of club or stick.[10] Given the strong medieval trade connections between south-east England and the County of Flanders when the latter belonged to the Duchy of Burgundy, the name may have been derived from the Middle Dutch (in use in Flanders at the time) "krick"(-e), meaning a stick (crook).[10] Another possible source is the Middle Dutch word "krickstoel", meaning a long low stool used for kneeling in church and which resembled the long low wicket with two stumps used in early cricket.[11] According to Heiner Gillmeister, a European language expert of Bonn University, "cricket" derives from the Middle Dutch phrase for hockey, met de (krik ket)sen (i.e., "with the stick chase").[12] Gillmeister has suggested that not only the name but also the sport itself may be of Flemish origin.[12]
Some species, such as Gryllus assimilis, take off, fly, and land efficiently and well, while other species are clumsy fliers.[1] In some species, the hind wings are shed, leaving wing stumps, usually after dispersal of the insect by flight. In other species, they may be pulled off and consumed by the cricket itself or by another individual, probably providing a nutritional boost.[12]
Cricket has had a broad impact on popular culture, both in the Commonwealth of Nations and elsewhere. It has, for example, influenced the lexicon of these nations, especially the English language, with various phrases such as "that's not cricket" (that's unfair), "had a good innings" (lived a long life) and "sticky wicket". "On a sticky wicket" (aka "sticky dog" or "glue pot")[128] is a metaphor[129] used to describe a difficult circumstance. It originated as a term for difficult batting conditions in cricket, caused by a damp and soft pitch.[130]
Meanwhile, the British Empire had been instrumental in spreading the game overseas and by the middle of the 19th century it had become well established in Australia, the Caribbean, India, New Zealand, North America and South Africa.[42] In 1844, the first-ever international match took place between the United States and Canada.[43] In 1859, a team of English players went to North America on the first overseas tour.[44]
Crickets are kept as pets and are considered good luck in some countries; in China, they are sometimes kept in cages or in hollowed-out gourds specially created in novel shapes.[49] The practice was common in Japan for thousands of years; it peaked in the 19th century, though crickets are still sold at pet shops.[50] It is also common to have them as caged pets in some European countries, particularly in the Iberian Peninsula. Cricket fighting is a traditional Chinese pastime that dates back to the Tang dynasty (618–907). Originally an indulgence of emperors, cricket fighting later became popular among commoners.[51] The dominance and fighting ability of males does not depend on strength alone; it has been found that they become more aggressive after certain pre-fight experiences such as isolation, or when defending a refuge. Crickets forced to fly for a short while will afterwards fight for two to three times longer than they otherwise would.[52]
It is generally believed that cricket originated as a children's game in the south-eastern counties of England, sometime during the medieval period.[3] Although there are claims for prior dates, the earliest definite reference to cricket being played comes from evidence given at a court case in Guildford on Monday, 17 January 1597 (Julian calendar; equating to 30 January 1598 in the Gregorian calendar). The case concerned ownership of a certain plot of land and the court heard the testimony of a 59-year-old coroner, John Derrick, who gave witness that:[5][6][7]

More than 900 species of crickets are described; the Gryllidae are distributed all around the world except at latitudes 55° or higher, with the greatest diversity being in the tropics. They occur in varied habitats from grassland, bushes, and forests to marshes, beaches, and caves. Crickets are mainly nocturnal, and are best known for the loud, persistent, chirping song of males trying to attract females, although some species are mute. The singing species have good hearing, via the tympana on the tibiae of the front legs.

Females are generally attracted to males by their calls, though in nonstridulatory species, some other mechanism must be involved. After the pair has made antennal contact, a courtship period may occur during which the character of the call changes. The female mounts the male and a single spermatophore is transferred to the external genitalia of the female. Sperm flows from this into the female's oviduct over a period of a few minutes or up to an hour, depending on species. After copulation, the female may remove or eat the spermatophore; males may attempt to prevent this with various ritualised behaviours. The female may mate on several occasions with different males.[18]

Crickets are relatively defenceless, soft-bodied insects. Most species are nocturnal and spend the day hidden in cracks, under bark, inside curling leaves, under stones or fallen logs, in leaf litter, or in the cracks in the ground that develop in dry weather. Some excavate their own shallow holes in rotting wood or underground and fold in their antennae to conceal their presence. Some of these burrows are temporary shelters, used for a single day, but others serve as more permanent residences and places for mating and laying eggs. Crickets burrow by loosening the soil with the mandibles and then carrying it with the limbs, flicking it backwards with the hind legs or pushing it with the head.[4]
Gryllus firmus exhibits wing polymorphism; some individuals have fully functional, long hind wings and others have short wings and cannot fly. The short-winged females have smaller flight muscles, greater ovarian development, and produce more eggs, so the polymorphism adapts the cricket for either dispersal or reproduction. In some long-winged individuals, the flight muscles deteriorate during adulthood and the insect's reproductive capabilities improve.[13]
Crickets have many natural enemies and are subject to various pathogens and parasites. They are eaten by large numbers of vertebrate and invertebrate predators and their hard parts are often found during the examination of animal intestines.[4] Mediterranean house geckos (Hemidactylus turcicus) have learned that although a calling decorated cricket (Gryllodes supplicans) may be safely positioned in an out-of-reach burrow, female crickets attracted to the call can be intercepted and eaten.[17]
Crickets are found in many habitats. Members of several subfamilies are found in the upper tree canopy, in bushes, and among grasses and herbs. They also occur on the ground and in caves, and some are subterranean, excavating shallow or deep burrows. Some make home in rotting wood, and certain beach-dwelling species can run and jump over the surface of water.[1]
Previous versions of the Spirit identified actions that were deemed contrary (for example, appealing knowing that the batsman is not out) but all specifics are now covered in the Laws of Cricket, the relevant governing playing regulations and disciplinary codes, or left to the judgement of the umpires, captains, their clubs and governing bodies. The terse expression of the Spirit of Cricket now avoids the diversity of cultural conventions that exist in the detail of sportsmanship – or its absence.
Other defensive strategies are the use of camouflage, fleeing, and aggression. Some species have adopted colourings, shapes, and patterns that make it difficult for predators that hunt by sight to detect them. They tend to be dull shades of brown, grey, and green that blend into their background, and desert species tend to be pale. Some species can fly, but the mode of flight tends to be clumsy, so the most usual response to danger is to scuttle away to find a hiding place.[4]
The wicket-keeper (sometimes called simply the "keeper") is a specialist fielder subject to various rules within the Laws about his equipment and demeanour. He is the only member of the fielding side who can effect a stumping and is the only one permitted to wear gloves and external leg guards.[77] Depending on their primary skills, the other ten players in the team tend to be classified as specialist batsmen or specialist bowlers. Generally, a team will include five or six specialist batsmen and four or five specialist bowlers, plus the wicket-keeper.[78][79]
Before a match begins, the team captains (who are also players) toss a coin to decide which team will bat first and so take the first innings.[63] Innings is the term used for each phase of play in the match.[63] In each innings, one team bats, attempting to score runs, while the other team bowls and fields the ball, attempting to restrict the scoring and dismiss the batsmen.[64][65] When the first innings ends, the teams change roles; there can be two to four innings depending upon the type of match. A match with four scheduled innings is played over three to five days; a match with two scheduled innings is usually completed in a single day.[63] During an innings, all eleven members of the fielding team take the field, but usually only two members of the batting team are on the field at any given time. The exception to this is if a batsman has any type of illness or injury restricting his or her ability to run, in this case the batsman is allowed "A Runner" who can run between the wickets when the batsman hits a scoring run or runs.[63] The order of batsmen is usually announced just before the match, but it can be varied.[58]
Crickets have a cosmopolitan distribution, being found in all parts of the world with the exception of cold regions at latitudes higher than about 55° North and South. They have colonised many large and small islands, sometimes flying over the sea to reach these locations, or perhaps conveyed on floating timber or by human activity. The greatest diversity occurs in tropical locations, such as in Malaysia, where 88 species were heard chirping from a single location near Kuala Lumpur. A greater number than this could have been present because some species are mute.[1]