The game underwent major development in the 18th century to become England's national sport. Its success was underwritten by the twin necessities of patronage and betting. Cricket was prominent in London as early as 1707 and, in the middle years of the century, large crowds flocked to matches on the Artillery Ground in Finsbury. The single wicket form of the sport attracted huge crowds and wagers to match, its popularity peaking in the 1748 season. Bowling underwent an evolution around 1760 when bowlers began to pitch the ball instead of rolling or skimming it towards the batsman. This caused a revolution in bat design because, to deal with the bouncing ball, it was necessary to introduce the modern straight bat in place of the old "hockey stick" shape.
Previous versions of the Spirit identified actions that were deemed contrary (for example, appealing knowing that the batsman is not out) but all specifics are now covered in the Laws of Cricket, the relevant governing playing regulations and disciplinary codes, or left to the judgement of the umpires, captains, their clubs and governing bodies. The terse expression of the Spirit of Cricket now avoids the diversity of cultural conventions that exist in the detail of sportsmanship – or its absence.
Some species of cricket are polyandrous. In Gryllus bimaculatus, the females select and mate with multiple viable sperm donors, preferring novel mates. Female Teleogryllus oceanicus crickets from natural populations similarly mate and store sperm from multiple males. Female crickets exert a postcopulatory fertilization bias in favour of unrelated males to avoid the genetic consequences of inbreeding. Fertilization bias depends on the control of sperm transport to the sperm storage organs. The inhibition of sperm storage by female crickets can act as a form of cryptic female choice to avoid the severe negative effects of inbreeding. Controlled-breeding experiments with the cricket Gryllus firmus demonstrated inbreeding depression, as nymphal weight and early fecundity declined substantially over the generations' this was caused as expected by an increased frequency of homozygous combinations of deleterious recessive alleles.
Cricket is one of many games in the "club ball" sphere that basically involve hitting a ball with a hand-held implement; others include baseball, golf, hockey, tennis, squash, badminton and table tennis. In cricket's case, a key difference is the existence of a solid target structure, the wicket (originally, it is thought, a "wicket gate" through which sheep were herded), that the batsman must defend. The cricket historian Harry Altham identified three "groups" of "club ball" games: the "hockey group", in which the ball is driven to and fro between two targets (the goals); the "golf group", in which the ball is driven towards an undefended target (the hole); and the "cricket group", in which "the ball is aimed at a mark (the wicket) and driven away from it".
Red parasitic mites sometimes attach themselves to the dorsal region of crickets and may greatly affect them. The horsehair worm Paragordius varius is an internal parasite and can control the behaviour of its cricket host and cause it to enter water, where the parasite continues its lifecycle and the cricket likely drowns. The larvae of the sarcophagid fly Sarcophaga kellyi develop inside the body cavity of field crickets. Female parasitic wasps of Rhopalosoma lay their eggs on crickets, and their developing larvae gradually devour their hosts. Other wasps in the family Scelionidae are egg parasitoids, seeking out batches of eggs laid by crickets in plant tissues in which to insert their eggs.
Crickets are small to medium-sized insects with mostly cylindrical, somewhat vertically flattened bodies. The head is spherical with long slender antennae arising from cone-shaped scapes (first segments) and just behind these are two large compound eyes. On the forehead are three ocelli (simple eyes). The pronotum (first thoracic segment) is trapezoidal in shape, robust, and well-sclerotinized. It is smooth and has neither dorsal or lateral keels (ridges).
In 1611, the year Cotgrave's dictionary was published, ecclesiastical court records at Sidlesham in Sussex state that two parishioners, Bartholomew Wyatt and Richard Latter, failed to attend church on Easter Sunday because they were playing cricket. They were fined 12d each and ordered to do penance. This is the earliest mention of adult participation in cricket and it was around the same time that the earliest known organised inter-parish or village match was played – at Chevening, Kent. In 1624, a player called Jasper Vinall died after he was accidentally struck on the head during a match between two parish teams in Sussex.
The fly Ormia ochracea has very acute hearing and targets calling male crickets. It locates its prey by ear and then lays its eggs nearby. The developing larvae burrow inside any crickets with which they come in contact and in the course of a week or so, devour what remains of the host before pupating. In Florida, the parasitic flies were only present in the autumn, and at that time of year, the males sang less but for longer periods. A trade-off exists for the male between attracting females and being parasitized.
There are various formats ranging from Twenty20, played over a few hours with each team batting for a single innings of 20 overs, to Test matches, played over five days with unlimited overs and the teams each batting for two innings of unlimited length. Traditionally cricketers play in all-white kit, but in limited overs cricket they wear club or team colours. In addition to the basic kit, some players wear protective gear to prevent injury caused by the ball, which is a hard, solid spheroid made of compressed leather with a slightly raised sewn seam enclosing a cork core which is layered with tightly wound string.
At the grassroots level, local club cricket is essentially an amateur pastime for those involved but still usually involves teams playing in competitions at weekends or in the evening. Schools cricket, first known in southern England in the 17th century, has a similar scenario and both are widely played in the countries where cricket is popular. Although there can be variations in game format, compared with professional cricket, the Laws are always observed and club/school matches are therefore formal and competitive events. The sport has numerous informal variants such as French cricket.
The innings (ending with 's' in both singular and plural form) is the term used for each phase of play during a match. Depending on the type of match being played, each team has either one or two innings. Sometimes all eleven members of the batting side take a turn to bat but, for various reasons, an innings can end before they have all done so. The innings terminates if the batting team is "all out", a term defined by the Laws: "at the fall of a wicket or the retirement of a batsman, further balls remain to be bowled but no further batsman is available to come in". In this situation, one of the batsmen has not been dismissed and is termed not out; this is because he has no partners left and there must always be two active batsmen while the innings is in progress.
A phylogenetic study by Jost & Shaw in 2006 using sequences from 18S, 28S, and 16S rRNA supported the monophyly of Ensifera. Most ensiferan families were also found to be monophyletic, and the superfamily Gryllacridoidea was found to include Stenopelmatidae, Anostostomatidae, Gryllacrididae and Lezina. Schizodactylidae and Grylloidea were shown to be sister taxa, and Rhaphidophoridae and Tettigoniidae were found to be more closely related to Grylloidea than had previously been thought. The authors stated that "a high degree of conflict exists between the molecular and morphological data, possibly indicating that much homoplasy is present in Ensifera, particularly in acoustic structures." They considered that tegmen stridulation and tibial tympanae are ancestral to Ensifera and have been lost on multiple occasions, especially within the Gryllidae.
The main objective of each team is to score more runs than their opponents but, in some forms of cricket, it is also necessary to dismiss all of the opposition batsmen in their final innings in order to win the match, which would otherwise be drawn. If the team batting last is all out having scored fewer runs than their opponents, they are said to have "lost by n runs" (where n is the difference between the aggregate number of runs scored by the teams). If the team that bats last scores enough runs to win, it is said to have "won by n wickets", where n is the number of wickets left to fall. For example, a team that passes its opponents' total having lost six wickets (i.e., six of their batsmen have been dismissed) have won the match "by four wickets".